Species List

** Reported locations based on a limited literature search. Codes indicate presence reported but absence of a code should not be interpreted as a species absence.

* (Dominant) based on:

• Relationships among zoobenthos, sediments, and organic matter in littoral zones of western Lake Erie and Saginaw Bay. 1983. Richard A. Cole and Diana L. Weigmann. JGLR 9(4)568-581.
• Disruption of the benthic community in Lake Ontario. 2003. S.J. Lozano and T.F. Nalepa. in State of Lake Ontario (SOLO) -- Past, Present and Future. M Munawar (ed). Ecovision World Monograph Series. Aquatic Ecosystem Health and Management Society.

S (Lake Superior) based on:

• Vertical distribution of profundal benthos in Lake Superior Sediments. 1982. Ann S. Heuschele. JGLR 8(4)603-613.
• Wave-zone macrobenthos of the exposed Canadian shores of the St. Lawrence Great Lakes. 1978. D.R. Barton and B.N. Hynes. JGLR 4(1)27-45.
• Benthic Community Structure and Composition among rocky habitats in the Great Lakes and Keuka Lake, NY. 1987. Michael H. Winnell and David J. Jude. JGLR 13(1)3-17.
• Lake Superior: an invasion coldspot? 2003. Igor A. Grigorovich, Alexei V. Korniushin, Derek K. Gray, Ian C. Duggan, Robert I. Colautti& Hugh J. MacIsaac. Hydrobiologia 499: 191–210.

M (Lake Michigan) based on combination of:

• The Nearshore Benthic Invertebrate Community of Southern Lake Michigan and its Response to Beach Nourishment. 2004. Eric L. Garza and Richard L. Whitman. J. Great Lakes Res. 30(1):114–122.

H (Lake Huron) based on:

• Abundance, Biomass, and Species Composition of Benthic Macroinvertebrate Populations in Saginaw Bay, Lake Huron, 1987-96. Thomas Nalepa, David Fanslow, Margaret Lansing, Gregory Lang, Mark Ford, Gerald Gostenik, and David Hartson. NOAA Technical Memorandum GLERL-122.
• Relationships among zoobenthos, sediments, and organic matter in littoral zones of western Lake Erie and Saginaw Bay. 1983. Richard A. Cole and Diana L. Weigmann. JGLR 9(4)568-581.
• Wave-zone macrobenthos of the exposed Canadian shores of the St. Lawrence Great Lakes. 1978. D.R. Barton and B.N. Hynes. JGLR 4(1)27-45.
• Benthic Community Structure and Composition among rocky habitats in the Great Lakes and Keuka Lake, NY. 1987. Michael H. Winnell and David J. Jude. JGLR 13(1)3-17.
• NOAA Great Lakes Nonindigenous Species List

C (St. Clair - Detroit River Corridor) based on:

• NOAA Great Lakes Nonindigenous Species List
• Moitessier's pea clam Pisidium moitessierianum (Bivalvia, Sphaeriidae): a cryptogenic mollusc in the Great Lakes. Igor A. Grigorovich, Alexei V. Korniushin, and Hugh J. MacIsaac. Hydrobiologia 435 (1-3): 153-165, September 2000
  
  

E (Lake Erie) based on:

• Relationships among zoobenthos, sediments, and organic matter in littoral zones of western Lake Erie and Saginaw Bay. 1983. Richard A. Cole and Diana L. Weigmann. JGLR 9(4)568-581.
• Wave-zone macrobenthos of the exposed Canadian shores of the St. Lawrence Great Lakes. 1978. D.R. Barton and B.N. Hynes. JGLR 4(1)27-45.
• Distribution of some common benthic invertebrates in nearshore Lake Erie, with emphasis on depth and type of substratus. 1988. David R. Barton. JGLR 14(1)34-43.
• Benthic and pelagic secondary production in Lake Erie after the invasion of Dreissena spp. with implications for fish production. 2000. Ora E. Johannsson, Ron Dermott, Donna M Graham, Julie A. Dahl, E. Scott Millard, Debra D. Myles, and Jennifer LeBlanc. JGLR 26(1)31-54.
• Changes in the benthic macroinvertebrate community of the Cleveland Harbor area of Lake Erie from 1978 to 1989. 1993. Kenneth A. Krieger and Lloyd S. Ross. JGLR 19(2) 237-249.
• Changes to the deepwater benthos of eastern Lake Erie since the invasion of Dreissena 1979-1993. 1997. R Dermott and D Kerec. CJFAS 54: 922-930.
• NOAA Great Lakes Nonindigenous Species List

O (Lake Ontario) based on:

• Disruption of the benthic community in Lake Ontario. 2003. S.J. Lozano and T.F. Nalepa. in State of Lake Ontario (SOLO) -- Past, Present and Future. M Munawar (ed). Ecovision World Monograph Series. Aquatic Ecosystem Health and Management Society.
• Wave-zone macrobenthos of the exposed Canadian shores of the St. Lawrence Great Lakes. 1978. D.R. Barton and B.N. Hynes. JGLR 4(1)27-45.
• Factors influencing changes in the nearshore benthic community on the Canadian site of Lake Ontario. 2000. Bruce W. Kilgour, Robert C. Bailey, and E Todd Howell. JGLR 26(3)272-286.
• Benthic macroinvertebrate communities of southwestern Lake Ontario following invasion of Dreissena. 1994. Timothy W. Stewart and James M. Haynes. JGLR 20(2)479-493.
• Benthic macroinvertebrate communities in southwestern Lake Ontario following invasion of Dreissena: Continuing change. 1999. James M. Haynes, Timothy W. Stewart, and George E. Cook. JGLR 25(4)828-838.
• NOAA Great Lakes Nonindigenous Species List
• Changes to the deepwater benthos of eastern Lake Erie since the invasion of Dreissena 1979-1993. 1997. R Dermott and D Kerec. CJFAS 54: 922-930.

X (Great Lakes coastal wetlands) based on:

• The ecology of invertebrates in Great Lakes coastal wetlands: current knowledge and research needs. 1992. Kenneth A. Krieger. JGLR 18(4)634-650.
• Burton, T.M., D.G. Uzarski and J.A. Genet. 2004.  Invertebrate habitat use in relation to fetch and plant zonation in northern Lake Huron coastal wetlands.  Aquatic Ecosystem Health & Management. 7(2): 249-267

W (wave zone = 0-2m) based on: Wave-zone macrobenthos of the exposed Canadian shores of the St. Lawrence Great Lakes. 1978. D.R. Barton and B.N. Hynes. JGLR 4(1)27-45.

R (rocky habitats) based on: Benthic Community Structure and Composition among rocky habitats in the Great Lakes and Keuka Lake, NY. 1987. Michael H. Winnell and David J. Jude. JGLR 13(1)3-17.

L (littoral) based on: Relationships among zoobenthos, sediments, and organic matter in littoral zones of western Lake Erie and Saginaw Bay. 1983. Richard A. Cole and Diana L. Weigmann. JGLR 9(4)568-581.

N (nearshore) based on:

• The Nearshore Benthic Invertebrate Community of Southern Lake Michigan and its Response to Beach Nourishment. 2004. Eric L. Garza and Richard L. Whitman. J. Great Lakes Res. 30(1):114–122.
• Distribution of some common benthic invertebrates in nearshore Lake Erie, with emphasis on depth and type of substratus. 1988. David R. Barton. JGLR 14(1)34-43.
• Factors influencing changes in the nearshore benthic community on the Canadian site of Lake Ontario. 2000. Bruce W. Kilgour, Robert C. Bailey, and E Todd Howell. JGLR 26(3)272-286.

1 = 14-30m based on:

• Disruption of the benthic community in Lake Ontario. 2003. S.J. Lozano and T.F. Nalepa. in State of Lake Ontario (SOLO) -- Past, Present and Future. M Munawar (ed). Ecovision World Monograph Series. Aquatic Ecosystem Health and Management Society.
• Factors influencing changes in the nearshore benthic community on the Canadian site of Lake Ontario. 2000. Bruce W. Kilgour, Robert C. Bailey, and E Todd Howell. JGLR 26(3)272-286.

2 = 31-50m based on:Disruption of the benthic community in Lake Ontario. 2003. S.J. Lozano and T.F. Nalepa. in State of Lake Ontario (SOLO) -- Past, Present and Future. M Munawar (ed). Ecovision World Monograph Series. Aquatic Ecosystem Health and Management Society.

3 = 51-70m based on:Disruption of the benthic community in Lake Ontario. 2003. S.J. Lozano and T.F. Nalepa. in State of Lake Ontario (SOLO) -- Past, Present and Future. M Munawar (ed). Ecovision World Monograph Series. Aquatic Ecosystem Health and Management Society.

4 = 71-90m based on:Disruption of the benthic community in Lake Ontario. 2003. S.J. Lozano and T.F. Nalepa. in State of Lake Ontario (SOLO) -- Past, Present and Future. M Munawar (ed). Ecovision World Monograph Series. Aquatic Ecosystem Health and Management Society.

5 = >90m based on:Disruption of the benthic community in Lake Ontario. 2003. S.J. Lozano and T.F. Nalepa. in State of Lake Ontario (SOLO) -- Past, Present and Future. M Munawar (ed). Ecovision World Monograph Series. Aquatic Ecosystem Health and Management Society.

Photos Courtesy of:

Field guide to Freshwater Mussels of the Midwest

Freshwater molluscan shells - Save the Clams

USGS - Gainesville

Conchology Inc.

Malakologie