Ph.D., Oregon State University, 1973, Biological Oceanography
M.S., University of Wisconsin, Madison, 1968, Zoology
B.S., Michigan Technological University, 1966, Biological Sciences
Interests are food web interactions, particularly feeding mechanisms and life cycle strategies
of plankton and benthos; and spatial distribution and interactions of all food web components.
Current research projects include: zooplankton feeding and mechanisms; dreissenid mussel
feeding behavior, nutrient cycling, and ecosystem effects; long-term trends in zooplankton
populations; impacts of the invasion of predatory cercopagids (Bythotrephes and
Cercopagis) on Great Lakes food webs; and changes in spatial structure and function
of Great Lakes food webs as driven by stressors such as hypoxia and invasive species.